the group of wrestlers such as chāṇūra, mushtika et al —

 chāṇūra and mushtika —

 wrestlers such as chāṇūra et al —

 the wrestlers, —

wrestlers chanuran and mushtikan —

wrestlers like Chanura and Mushtika —

wrestlers —

wrestlers (chāṇūra and mushtika) —

and who destroyed the wrestlers (chāṇūra-mushtika) —

and also the wrestlers —